Eastern Sphecid Related Wasps


Working list of the possible Eastern Sphecid wasps with new families, subfamilies, tribes and subtribes noted. (See this paper from the wasp man, Pulawski, with some reordering of the family groups. I am working on shuffling the list around based on this update.)


Otherwise taken from combined sources including the internet and the impressive but slightly outdated Sphecid Wasps of the World from Bohart and Menke. Ranges are spotty with states and regions listed for known occurrences. These are now updated to literature references from the Catalog update mentioned below. I have no lists of confirmed Arkansas species as of yet. Species with AR in blue with a numeral are in the collection at UA Fayetteville and have the number of counties represented by the numeral. Species will be removed from the list when I have information narrowing the ranges outside of the east. "Nea" refers to number of species listed for all of North America at the excellent Nearctica insect list.


And here is the updated Sphecid Wasp Catalog from the California Academy of Sciences (covers the entire globe but it is more UTD). Presently there are about 1400 species in North America. Habits and identifying characteristics given where known. Prey choice of species and groups marked in red where known from various sources.







Ampulex (2 Nea) (Uses cockroaches, nymphal, one per cell, pulls prey by antennae bases; nests in twigs, stems and borings; Krombein traps; partitions of leaves and grass)

            canaliculata Say 1823 (AR CT MI MO NC PA WV) (July to Sept in NC)

            ferruginea Bradley 1934 (OK TX)




Dolichurus (1 Nea) (Uses cockroaches; nests in stems and crevices)

            greenei Rohwer 1916 (FL MI NC VA)






Podalonia (19 Nea) (Uses Noctuid larvae from soil, searches over soil and then digs; captures prey first, stashes and dig sometimes twenty feet away; caterpillars carried venter up by mandibles over ground; late summer and early fall females may be found in mass clusters in crevices)

            luctuosa (F. Smith) 1856 (CO MI NE NC TX UT)

(1 Noctuid per cell) (body black with a faint bluish tinge)

            valida (Cresson) 1865 (CA CO NE TX)

            violaceipennis (Lepeletier) 1845 (FL MA MD MI NC ND PA)

(abdomen partly reddish)


Eremnophila (1 Nea) (Uses large caterpillars mostly of Notodontidae; carries venter up over ground by mandibles)

            aureonotata (Cameron) 1888 (AR4 FL MD MI NY NC TX WV)


Ammophila (62 Nea) Caterpillar hunters (Uses Lepidopterous and Hymenopterous [sawfly] caterpillars; dig nests first; carries prey head first, venter up with mandibles and sometimes front pair of legs; often flies with prey)

            evansi Menke 1964 (CT MI NC NY)

(uses loamy soil among vegetation)

            extremitata Cresson 1865 (AR1 CA CT IL ID NE)

            gracilis Lepeletier 1845 (MD TX)

            harti (Fernald) 1931 (CO IL MI MO NE NY WY)

(sand dunes and blowouts; 7 familes of Geometrid; 5-12 items per cell)

            kennedyi (Murray) 1865 (CA CO CT IL MA MI NE NM NY TX) (close to urnaria)

            leoparda (Fernald) 1934 (GA MI MN) (uncommon)

            nigricans Dahlbom 1843 (AR6 IL KS MI NE SC NC NY PA) (sandy clay soils)

            pictipennis Walsh 1869 (AR1 CO IL IA KS MD MI MO NC NJ NY TX)

(shallow unicellular nests; uses Noctuid larvae in Michigan)

            placida F. Smith 1856 (CA FL MA MI NY)

            procera Dahlbom 1843 (AR3 AZ CA CO CT FL ID IL IA KS MD MI MO NE NC TX MN UT)

(large, two stripes and a dash of silver on thorax)

            urnaria Dahlbom 1843 (AR3 CO CT FL IL MD MI MN NE NY PA SC VA TX WV)

(waste areas and gravel pits; uses Geometrids and Noctuids in Michigan)

            varipes Cresson 1865 (IA NE NM TX)




Chlorion (2 Nea) (Uses Gryllids, up to seven per nest)

            aerarium Patton 1879 2nd (AR5 CT FL KS MA MI NC TX)

(Gryllidae, 2-9 per cell)

            cyaneum Dahlbom 1843 (CO GA IL MI MO NE NC TX)




Subtribe Sceliphina


Chalybion (2 Nea) (Uses spiders; nests in holes in wood; sometimes takes over mud nests of others; carries water to dirt to make usable mud)

            californicum (de Saussure) 1763 (AR6 CA CO CT FL KS MI NM NE NC OH OK PA SC TX UT WV)

            zimmermanni zimmermanni Dahlbom 1843 (AZ IN NC SC TX) (seals nest with white material)


Sceliphron (2 Nea) (Uses spiders, favors Argiopidae; makes globular mud nests)

            caementarium (Drury) 1770 (AR8 CA CO CT FL LA MD MI NE NC NY PA TX UT VA)

(Argiopidae, 6-15 per cell)


Subtribe Podiina


Podium (3 Nea) (Uses Cockroaches; nests in wood chambers or bark; prey is held in mandibles)

            fulvipes Cresson 1865 (FL only)

            krombeini Bohart and Menke 1963 (TX only)

            luctuosum F. Smith 1856 (DC FL LA MD MI NC NY TX) (Krombein with only one nest in his entire study)

            rufipes Fabricius 1804 (FL MI MO NC TN TX) (complex nest closure; 3-14 per cell)


Sphecini (Use Tettigonids)


Subtribe Sphecina


Isodontia (6 Nea) (Uses Gryllids or Tettigonidae; nests in hollow stems, rolled leaves; cuts grass for closures)

            apicalis (F. Smith) 1856 (AR1 FL IL NC TX) (Tettigonids)

            auripes (Fernald) 1906 2nd (AR FL MA MI MD MO NC PA SC TN TX)

(grass) (Gryllids 9-18 per cell, common brood chamber) (three toothed mandible, yellow legs)

            azteca (de Saussure) 1867 (AR1 FL MI NC)

            elegans (F. Smith) 1856 (CA NE TX)

            exornata Fernald 1903 (FL GA NC)

            mexicana (de Saussure) 1867 (AR2 FL GA IL KS MI MO NE NY NC PA TN TX)

(grass) (Gryllids and Tettigonids, up to 21 per cell) (known to use Pitcher Plants for nests in SE)

            philadelphica (Lepeletier de Saint Fargeau) 1845 (CO KS MA NC PA TX) (Tettigonids)


Sphex (13 Nea) (Use one species of katydid per wasp species; ground nesters, gregarious, repeat site users)

            ashmeadi (Fernald) 1906 (AZ CA TX)

            dorsalis Lepeletier de Saint Fargeau 1845 (AR1 FL TX)

            flavovestitus F. Smith 1856 (FL GA NC VA TX)

            habenus Say 1831 2nd (FL LA NC TX VA)

            ichneumoneus (Linnaeus) 1758 2nd (AR5 AZ CA FL ID IL KS LA MA MD MI MO NE NC NY OH TX UT)

(abdomen and legs red or reddish, wings subhyaline)

            jamaicensis (Drury) 1773 (FL only)

            lucae de Saussure 1867 (AZ CA NE TX)

            nudus Fernald 1903 (IL MD NC SC TN TX)

(bright orange-red legs beyond mid-femur on all legs)

            pensylvanicus Linnaeus 1763 2nd (AR2 CT FL IL KS MA MD MI NE NC NY TX )

(black legs and abdomen, wings blackish)

            texanus Cresson 1873 (TX only)


Subtribe Prionyxina


Palmodes (10 Nea) (Uses Tettagonids; all western except dimidiatus; carries prey holding one antennae by straddling and pulling over ground)

            dimidiatus (De Geer) 1773 (CA FL KS MI NC PA TX)

(temp nest closures, single provisions, nest excavated first)


Prionyx (7 Nea) (Uses only Acridids, obtains prey first, then stashes and digs nest, holds prey by antennae with mandibles)

            atratus  (Lepeletier) 1845 (AR1 CO KS MI MO MT NE ND NM NY NC TX UT WY)

(uses single grasshopper) (dark brown or black abdomen)

            parkeri Bohart & Menke 1963 (AR2 AZ CO NC TX WY) (red abdomen)

            thomae (Fabricius) 1775 (CA CO FL GA MA MI MS NM NC TX) (red abdomen)






Astata (14 Nea) (Uses Heteroptera, Pentatomids; very fast flight)

            bicolor Say 1823 (AR CA CT KS NC NM NE NY TX) (abdomen red)

            leuthstromi Ashmead 1897 (CO ID NY VA WV WI)

            nubecula Cresson 1865 (CA CO ID NY NE ND UT WV)

            occidentalis Cresson 1881 (AR ID NV TX VA WA)

(3-6 per cell) (very dark winged, species appears much blacker than unicolor)

            unicolor Say 1824 (CT IA MD NY NM NC TX VA) (abdomen black)


Dryudella (10 Nea, essentially western genus) (Uses Heteroptera)

            pernix F. Parker 1969 (TX)







Alysson (8 Nea) (Uses Homoptera - leafhoppers; nests in moist cool sites, sandy soil, gregarious)

            conicus Provancher 1889 (MD NY PA)

            guignardi Provancher 1887 (IL NM QU)

            melleus Say 1837 (AR2 FL IN MA MD MO NE NY NC TN TX VA) (3-23 per cell)

            oppositus Say 1837 (IN KS MD NY NC)

            radiatus W. Fox 1894 (NE NC WY)

            striatus W. Fox 1894 (DC)

            triangulifera Provancher 1887 (CO IA KY MD NE NC NY)


Didineis (7 Nea) (Uses Homoptera and possibly Fulgorids)

            dilata Malloch and Rohwer 1930 (ONT WI)

            latimana Malloch and Rohwer 1930 (MD)

            texana (Cresson) 1873 (AR1 FL MD MO IA NC)


Bembicini (Fly hunters)


Subtribe Bembicina


Bembix (19 Nea) (Serial feeders, Bembix females sometimes steal from each other)

            americana spinolae Lepeletier (AR5 FL IL MA MD MO NC NY OH PA TX)

(favors Muscidae) (fasciae on tergites white, with second tergite like the third and fourth, none of the fasciae greatly attenuated medially, clypeus not marked with black)

            belfragei Cresson 1873 (GA KS LA MI NE NC TX WI)

(fasciae on all tergites interrupted medially, ultimate tergite strongly wrinkled)

            cinerea Handlirsch 1893 (FL GA NC)

(creamy white abdominal markings)

            hinei Parker 1917 (TX LA)

(closely related to cinerea, but in this species tarsi are invariably yellow and in cinerea all or mostly black, also mandibles are black on cinerea and yellow on this species)

            nubilipennis Cresson 1873 (AR1 CO IL IN IA KS MO NE NM OK TN TX)

(wings of female infuscated in medial portion, base and apex clear, male has lateral processes on the sixth sternite, fasciae yellow, clypeus wholly yellow)

            pallidipicta F. Smith 1873 (CO FL IA KS LA NJ NY NC NE NY OH OR)

(sand) (17-22 mm species, no trace of discal scutal marks, maculations white and broad, fifth tergite fasciae continuous, anterior metatarsus with seven spines, sixth tergite black)

            texana Cresson 1873 (AR2 CO FL GA IL LA NC NM TX)

(favors Tabanidae) (maculations white, wings almost hyaline, fasciae on the second tergite incloses either completely or incompletely a pair of black discal spots, black basal spots on the female clypeus)


Bicyrtes (8 Nea) (Uses Pentatomids,stinkbugs)

            capnoptera (Handlirsch) 1889 (AR3 AZ CA GA KY LA NE NM NC TX UT)

(legs black and ferruginous, maculations deep yellow, fasciae present on metanotum and fasciae on tergites relatively broad with sixth tergite black in female)

            fodiens (Handlirsch) 1889 (AR3 GA IN KS LA MI MS MO NE NC TX VA)

(fasciae on tergites narrow and ultimate tergite black, posterior coxa toothed on inner margin)

            insidiatrix (Handlirsch) 1889 (KY TX)

(flagellum and legs for the most part ferruginous, apex of ultimate tergite ferruginous, fasciae on first tergite broad and best developed, mesopleura black)

            quadrifasciata (Say) 1824 (AL AR8 CO CT FL GA IL IN IA LA MI MO NY NC OH PA SC TX VA WI)

(favors Coreidae; 4-14 per cell) (18mm wasp) (fasciae on fifth tergite much more widely interrupted than on the first, discal marks on scutum lacking or reduced, wing not clouded, ultimate tergite black, mesopleura maculated with yellow)

            ventralis (Say) 1824 (CA CO KS MI NE NM NV NY NC PA TX UT)

(Holds prey upside down by middle legs) (metanotom with yellow spots, ultimate tergite black, flagellum black, fasciae on first tergite narrow, mesopleura black)

            viduatus (Handlirsch) 1889 (AZ IN KS TX)

(anterior wing clouded at first cubital cell, ultimate tergite black, mesopleura maculated with yellow)


Glenostictia (19 Nea) (Progressive provisioners)

            pictifrons (F. Smith) 1856 (AR4 CA CO GA IN MI MO NE NM NC PA TX VA)

(Uses Bombylid flies) (discal marks on scutum consisting of a pair of irregular spots, fasciae on first tergite deeply emarginate)


Microbembex (8 Nea)

            evansi R. Bohart 1993 (TX)

            hirsuta J. Parker 1917 (TX)

(pubescence on head, thorax and propodeum unusually long and dense)

            monodonta (Say) 1824 (AR4 CO CT KS MO NE NC ND PA TX VA WY)



            eos Ohl 2005 (FL only)

            femorata (W. Fox) 1895 (FL TX)

(10 to 12 mm species, discal marks on scutum narrow and broken, fasciae on tergites rather narrow and minimally emarginate, unite with the fasciae on the sternites to form almost strict rings around the abdomen)


Stictia (3 Nea) (Uses Diptera esp. Horseflies)

            carolina (Fabricius) 1793 (AL AR3 FL GA IL KS LA MD NJ NC NM OK SC TX)

(sometimes hovers around horses for its prey, sandy riverside soil in my area)

            vivida (Handlirsch) 1890 (TX)

(labrum and clypeus wholly yellow, all fasciae interrupted but not broken into spots)


Stictiella (18 Nea)

            emarginata (Cresson) 1865 (CO FL GA IN KS MI NE ND WY)

            formosa (Cresson) 1873 (AR3 CO KS OK TX)

(favors Libytheidae; up to 11 per cell) (16 to 20 mm large species, discal marks on scutum two broad lines curved and approximated posteriorly)

            pulchella serrata (Handlirsch) 1890 (FL GA MI NC)

(Uses adult moths and butterflies) (fasciae on first tergite interrupted widely)


Subtribe Exeirina


Clitemnestra (1 Nea) (Uses Hemiptera; nests in sandy soil)

            bipunctata (Say) 1824 (CA CT FL MA MD NJ NY NC PA VA WV WY)


Subtribe Gorytina


Argogorytes (2 Nea)

            nigrifrons (F. Smith) 1856 (MD NC TX)


Gorytes (16 Nea)

            atricornis Packard 1867 (CT IL MA NE NY TX)

            canaliculatus Packard 1867 (CA CO MD NE NC NY PA WY)

            deceptor Krombein 1958 (TX WV)

            dorothyae Krombein 1950 (NC TX)

            dorothyae russeolus Krombein 1954 (FL)

            mcateei Krombein and Bohart 1962 (MD)

            simillimus F. Smith 1856 (DC ID IL NY NC PA VA WV)


Oryttus (5 Nea)

            gracilis (Patton) 1879 (CT NE ONT)

            gracilis arapaho (Pate) 1938 (CO TX)


Psammaletes (3 Nea)

            floridicus R. Bohart 2000 (FL)

            mexicanus (Cameron) 1890 (NY NC TX VA)


Pseudoplisus (19 Nea)

            apicalis (F. Smith) 1856 (GA NE NM)

            californicus R. Bohart 1969 (CA TX)

            divisus (F. Smith) (GA NE NC TX)

            phaleratus (Say) (AR4 AZ CO CT FL IL IN MD MA NE NJ NM NY NC PA TX)

            rufomaculatus (W. Fox) 1896 (NE TX)

            smithii (Cresson) 1880 (AR2 IL KS NE TX)

            smithii floridanus (W. Fox) 1890 (FL NE NC VA)

            tanythrix R. Bohart 1969 (OK)

            venustus (Cresson) 1865 (CO NM NC)


Hoplisoides (17 Nea) (Uses Homoptera)

            costalis (Cresson) 1873 (MO NC OH TX)

            denticulatus (Packard) 1867 (AR1 GA IL LA NC TX)

            floridicus R. Bohart 1968 (FL)

            nebulosus (Packard) 1867 (AR1 GA NE NJ NC ND TX VA)


Subtribe Handlirschiina


Sphecius (4 Nea)

            grandis (Say) 1823 (AR NV TX)

            hogardii (Latreille) 1806 (FL only)

            speciosus (Drury) 1773 (AR5 CO FL KS MD MO NE NY NC PA TX) (uses Cicadidae; 1-4 per cell)


Tanyoprymnus (1 Nea)

            moneduloides (Packard) 1867 (LA NM NC TX)


Subtribe Stizina


Bembecinus (6 Nea)

            nanus (Handlirsch) 1892 (CA DC NC)

(progressive feeders, favors leafhoppers and especially Dictyopharidae; 3 per cell)

            neglectus (Cresson) 1873 (KS NE TX)


Stizoides (2 Nea) (kleptoparasite of wasps in Prionyx which also take grasshoppers)

            renicinctus (Say) 1823 (AR1 CO ID KS NE NM NC TX UT)


Stizus (5 Nea)

            brevipennis Walsh 1869 (AR2 CO IL IN KS MD NE NC)


Nyssonini (50 NA spp) (Kleptoparasites of Gorytini and Larrini)


Epinysson (12 Nea) (Parasites of Hoplisoides)

            basilaris (Cresson) 1882 (GA NC ND)

            mellipes (Cresson) 1882 (CO NE NJ NC TX VA)

            opulentus (Gerstaecker) 1867 (CA GA MD NY NC TX VA)

            tramosericus (Viereck) 1904 (NY PA)

            tuberculatus (Handlirsch) 1887 (MD SC VA WI)


Hyponysson (2 Nea)

            raui (Rohwer) 1917 (MO)


Nysson (23 Nea) (Kleptoparasites of Gorytes)

            aequalis Patton 1879 (MA NE NC TX)

            aurinotus (Say) 1837 (GA IN KS NE TX)

            daeckei Viereck 1904 (NJ PA WY)

            gagates Bradley 1920 (BC PA)

            lateralis Packard 1867 (MD ND VA WV)

            plagiatus Cresson 1867 (IL IA QU)

            simplicicornis W. Fox 1896 (MI MO NE WV WY)

            subtilis W. Fox 1896 (IL TX WV)

            trichrus (Mickel) 1916 (NE NY TX)

Zanysson (3 Nea) (
Parasitic on Tachytes)

            plesius (Rohwer) 1921 (AZ LA NY NC)

            texanus (Cresson) 1873 (AR1 KS NM NC TX)






Subtribe Bothynostethina


Bothynostethus (1 Nea) (Uses Chrysomelids; remodels tunnels)

            distinctus W. Fox 1891 (CO FL KS NJ NC) (4-9 per cell)


Crabronini (Provision nest with various groups as below; impaled on the sting)


Subtribe Anacrabronina


Anacrabro (1 Nea) (Carries prey with mid and hind legs; nectars at Umbels, Composites)

            ocellatus Packard 1866 2nd (DC FL IL KS MD ME NE NY NC TX) (uses Miridae; 4-9 per cell)


Entomognathus (4 Nea)

            arenivagus Krombein 1963 (FL NC)

            lenapeorum Viereck 1904 (KS MD NJ PA VA)

            memorialis Banks 1921 (KS MD NJ VA) (uses Chrysomelids; 3-9 per cell)

            texanus Cresson 1887 (KS OK PA TX)


Subtribe Crabronina


Crabro (46 Nea) (Holds prey venter up, beneath body by middle legs)

            advena F. Smith 1835 (AR1 Extensive east of the Rockies) (Uses cluster flies)

(main species in the East with the male having a shield, a bifid mandibular apex and the foretarsomere simple, see vernalis and henrici, shield strawberry shaped with a few small spines on foremargin and darkened overall with white colored fingers from hindmargin and leg)

            aequalis W. Fox 1895 (Extensive East)

(unshielded males group)

            arcadiensis R. Miller 1976 (FL only)

(in the hilaris group with no shields)

            argusinus R. Bohart 1835 (Extensive Eastern and rare in the West to CA) (favors Ephydridae; 5-92 per cell)

(modified foretarsomere group, male with distinctive pectinate antenna segments, shield is arrowhead shaped with two tone look, leg and base light and most of pointed head dark with light spotting)

            atlanticus R. Miller 1976 (GA NC NJ)

(in the hilaris group with unshielded males)

            cingulatus (Packard) 1867 (AR1 IL KS NE NC WA, most of East)

(unshielded males)

            cognatus W. Fox 1895 (Everything west of ID UT AZ)

(unshielded males)

            cribellifer (Packard) 1867 (Extensive Eastern) (favors Asilidae, mainly Ommatius in one report)

(modified foretarsomere group, shield uniform grayish triangular, indented on hind margin)

            digitatus R. Bohart 1976 (IL MI MN NY OH ONT)

(modified foretarsomere group, shield shape like cribellifer, lighter overall with light hindmargin edge)

            henrici Krombein 1951 (DC NJ NY PA VA)

(shielded males with bifid mandibles and simple foretarsomeres, shield human heart shaped [the real ones] with a dark spot along distal furrow near leg, mostly shadowed otherwise with light spotting)

            hilaris F. Smith 1856 (Extensive Eastern)

(unshielded males)

            latipes F. Smith 1856 (Entire US, possibly the most common species of Crabro)

(modified foretarsomere, shield elongate with undulate rear border and very pointed end with small spines, hand shaped darkness centrally with encroaching pale triangles)

            monticola (Packard) 1867 (GA to ME, MI MN) (Nests in aggregations)

(modified foretarsomere group, shield bluntly pointed on tarsomere side making it triangular, mostly dark especially at tip, some linear mottling on proximal half subparallel to leg)

            peltista Kohl 1888 (AZ NC? TX)

(shielded males with simple mandible, Christmas-stocking shaped shield with toe pointing towards tarsomeres, very dark with two light stripes in proximal one-third, midbasitarsus of male flattened and expanded)

            rufibasis (Banks) 1921 (FL only)

(unshielded males)

            snowii W. Fox 1896 (Extensive East including KS)

(unshielded males)

            tenuiglossa (Packard) 1866 (Extensive East)

(shielded males with simple mandible, shield broadly rounded with irregular dark patch, no stripes)

            tenuis W. Fox 1895 (Extensive East and NW)

(modified foretarsomere group, shield blunt tipped, leg portion half light and half dark parallel to leg, posterior fourth zebra striped from hindmargin)

            tumidus (Packard) 1867 (AR Extensive Eastern including NE TX)

(shielded males with simple mandible, broadly rounded shield with three darkened stripes parallel to leg)

            vernalis (Packard) 1867 (North and NW south to IL NJ)

(shielded males with bifid mandibles and simple foretarsomeres, shield very rounded with 8 to 10 dark broken stripes perpendicular to leg)


Crossocerus (30 Nea) (Uses small Diptera; carries prey venter up by midlegs; nests in sandy or clay soil)

            annulipes (Lepeletier and Brulle) 1835 (MD NC WV) (Uses leafhoppers; 14-30 per cell; nests in stumps or wood)

            arnoldi Leclerq and Miller 2000 (VA)

            barbipes (Dahlbom) 1845 (CO NC SD TN WA WV WY)

            elongatulus (Vander Linden) 1829 (CT MA PA)

            harringtonii (W. Fox) 1895 (MD NM WV)

            impressifrons (F. Smith) 1856 (DC FL MA NE NC TX VA WV)

            krombeini Leclerq and Miller 2000 (NY)

            lentus (W. Fox) 1895 (FL IL KS MD NE NY NC PA VA WV)

            maculiclypeus (W. Fox) 1895 (KS NJ NC PA)

            minimus (Packard) 1867 (ME NC TX WV)

            nitidiventris (W. Fox) 1892 (IL MD NC PA VA)

            planifemur Krombein 1952 (MD NC WV)

            similis (W. Fox) 1895 (FL NJ SC TX VA WV)

            stictochilos Pate 1944 (MD NJ)

            tarsalis (W. Fox) 1895 (NY NC VA WV)

            tarsatus planipes (W. Fox) 1895 (KS MD PA)

            unicus (Patton) 1879 (CT PA)

            xanthochilos Pate 1944 (GA MD VA)


Ectemnius (27 Nea) (Uses adult Diptera; nests in decayed wood, twigs)

            arcuatus Say 1837 (IN ID CO NJ WA)

            atriceps Cresson 1865 (CO ID MD NM NC PA WA WV WY)

            cephalotes (Olivier) 1792 (MA)

            continuus Fabricius 1804 (AR3 MD NC PA ONT)

            decemmaculatus (Say) 1823 (AR1 CT FL GA MO MN NC ND PA TX UT)

            dilectus (Cresson) 1865 (CO CT NC ND ID IL UT WI)

            dives (Lepeletier and Brulle) 1835 (CO CT ID IL MN NM NY NC PA WY)

            lapidarius (Panzer) 1804 (AZ CA CO CT NY NC PA VA)

            maculosus (Gmelin) 1781 (CO CT IL KY MD MI NE ND NC ONT PA WA)

            paucimaculatus (Packard) 1866 (MD IL) (favors Ephydridae; 7-31 per cell)

            ruficornis (Zetterstedt) 1838 (CO KS NC WV)

            rufifemur Packard 1866 (IL NE TX)

            rufipes (Lepeletier and Brulle) 1835 (AL FL MO NE NC NY TX)

            rufipes ais (Pate) 1946 (FL)

            scaber (Lepeletier & Brulle) 1835 (AR1 NC PA)

            scaber rufescens Krombein 1954 (FL)

            stirpicola (Packard) 1866 (MD MO NY NC VA)

            trifasciatus Say 1824 (CT MN NY WY)


Enoplolindenius (2 Nea)

            clypeatus Rohwer 1911 (TX)

            robertsoni (Rohwer) 1920 (IL GA NE)


Lestica (3 Nea)

            confluenta (Say) 1837 (CO IN KS MA MO NE NY NC PA)

            producticollis (Packard) 1866 (NJ NC VA WV)


Lindenius (10 Nea)

            armaticeps (W. Fox) 1895 (CO NE NC NY TX VA) (uses Chloropidae; 3-15 per cell)

            buccadentis Mickel 1908 (AR NE NC NY VA WY)

            columbianus (Kohl) 1892 (AR3 NE NC NM NY PA WA WY)

            latifrons (W. Fox) 1895 (NC TX)

Rhopalum (7 Nea) (
Uses small Diptera; nests in twigs, reeds)

            atlanticum R. Bohart 1974 (GA)

            clavipes (Linnaeus) 1758 (NY)

            coarctatum (Scopoli) 1763 (WV)

            occidentalis (W. Fox) 1895 (NC NY NV)

            pedicellatum Packard 1867 (CO CT NC NY OH WI)

            rufigaster Packard 1867 (DC IL NC PA TX)


Moniaecera (5 Nea) (Uses Cicadellids; ground nester)

            abdominalis (W. Fox) 1895 (AZ GA KS NM TX)

            texana LeClerq 2003 (TX)




Subtribe Gastrosericina


Larropsis (24 Nea) (Uses Gryllids; males often found near gopher or rabbit holes)

            consimilis (W. Fox) 1894 (KS TX)

            discreta (W. Fox) 1894 (GA)

            distincta (F. Smith) 1856 (CT IA KS MA NE NC NY ONT PA)

            elegans Bohart and Bohart 1966 (TX only)

            filicornis Rohwer 1911 (NE TX)

            greenei Rohwer 1917 (NJ)

            sericea Bohart and Bohart 1966 (TX only)

            shappirioi Bohart and Bohart 1962 (DC)

            sparsa Bohart and Bohart 1966 (TX only)

            texensis Bohart and Bohart 1966 (TX only)


Tachysphex (76 Nea) (Uses Orthoptera; digs nest first, throwing soil out behind them in a stream; one-celled nests; grasps prey by antennae, legs assist)

            acutus (Patton) 1881 (CT KS MN NC NE NY WV WI)

(abdomen black or only apically red)

            antennatus W. Fox 1894 (CA FL KS NE NC TX WI)

(wings clear except apically, abdominal segments 1-3 reddish brown)

            apicalis W. Fox  1893 (AR1 CA CO CT FL KS NC NM ND SC TX WI)

(apex of abdomen red, dense silvery pubescence, F 9-11mm)

            belfragei (Cresson) 1873 (AR3 FL KS NE NC NY TX)

(tibia and tarsi reddish, segments 1, 2 and 3 reddish and apex of pygidium reddish, F 9-10mm)

            crassiformis Viereck 1906 (AR1 CA FL KS NC TX)

(abdomen reddish, distal tarsi reddish)

            krombeini Kurczewski 1971 (FL only)

            krombeiniellus Pulawski 1982 (FL only)

            laevifrons (L. Smith) 1856 (FL KS NC TX)

(abdomen black apically; tibia and femora black, clypeus entire)

            maurus Rohwer 1911 (TX only)

            mundus W. Fox 1894 (AR1 CO IL NC NE MT TX)

            pechumani Krombein 1938 (IN NJ)

(Acrididae, 1 per cell)

            pompiliformis (Panzer) 1805 (CO CT KS NE OR)

(abdomen with only first and second segments red; tibia and femora black)

            punctifrons (W. Fox) 1891 (CO FL MN MT NE NC TX)

            scopaeus Pulawski 1988 (TX only)

            similis Rohwer 1910 (AR2 FL MD MN NC NY SC TX VA WI)

            tarsatus (Say) 1823 (AR CA CO CT KS MA MN NE NY NJ NC PA TX WI WY)

(abdomen entirely red or red and black, 8-11 mm females)

            terminatus (F. Smith) 1856 (KS MA NE NM NC NY PA SC VA WI)

(uses Acrids, 3-7 per cell) (abdomen black with red apex)

            texanus (Cresson) 1873 (AR1 CA KS NE NC TX WI)

(wings clear, abdomen all red, 7.5-10 mm females)

            utina Pulawski 1988 (FL only)


Tachytes (34 Nea) (Uses Orthopterans; nests in preexisting holes in soil, rodent burrows, old or active Sphecius burrows; mounded soil at site; many-celled nests; pushes out soil with abdomen; prey flown to nest, carried by antennae)

            abdominalis (Say) 1823 (AR ID KS NM NC TX)

(favored Tettigidae and Melanopli in Williams 1913)(abdominal segments 1 and 2 or 1 to 3 ferruginous, abd with four pale golden fasciae, 9-10 mm)

            amazonus F. Smith 1856 (AR4 FL KS NE NC TX WI)

(wings nearly clear, pygidium bright golden, abd. variably red with golden fasciae)

            auricomans Bradley 1919 (East NC to FL, IL to East TX)

(scape dark, three light abdominal fasciae)

            aurulentus (Fabricius) 1804 (AR2 FL CO CT KS MD NE NC PA TX VA WI WY)

(scutum silvery to gold, pygidial dark coppery)

            badius Banks 1942 (TX only)

            birkmanni Rohwer 1909 (AZ TX)

(four bands on abdomen, 6-7 mm)

            chrysopyga obscurus Cresson 1873 (AR2 AZ KS MD MO NE NC TX)

(abdomen with four silvery fasciae, 1 to 3 black, F 13-14mm)

            columbiae W. Fox 1892 (MD to FL, IL TX)

(hind femur half to all red, propodeum golden haired)

            crassus Patton 1880 (CO CT DC MD NC NE WI)

(Uses Katydids) (abdomen black with three dorsal silver bands, hind femurs very hairy)

            cressoni Banks 1942 (TX only)

(slender spines on female hind tibia, 5 and 6 abdominal black)

            desertus Bohart 1994 (TX only)

(female hindfemur red)

            distinctus F Smith 1856 (AR6 CA CO GA KS NC NM PA TX UT)

(Uses mostly genus Melanopli in KS)

(legs mostly ferruginous, three silvery fasciae on abdomen, 15-22mm)

            floridanus Rohwer 1920 (FL GA KS NC TX)

(bright silvery pygidium)

            grisselli Bohart 1994 (FL only)

            guatamalensis Cameron (Eastern to TX KS)

(two dorsal abdominal bands on female)

            harpax Patton 1880 (CT KS NC NE PA WI)

(Uses genus Xiphidium in KS) (thorax with grayish pubescence, hind femur black)

            intermedius (Viereck) 1906 (DC FL KS MD NC NE NY TX)

(Uses Pygmy mole crickets; 2-6 per cell) (female abdominal 1 and 2 sometimes red)

            mergus W. Fox 1892 (AZ FL KS MO NJ NC TX WI)

(wings clear, F 9mm, mandible red or reddish)

            obductus W. Fox 1892 (FL KS MO NC NY TN TX)

(pygidium with fine bright golden pile, abd segments 1-5 brassy or silver fasciate, F 10.5mm)

            parvus W. Fox 1892 (AZ FL ID NC NE NJ NY)

(abdominal 1 to 3 red)

            pennsylvanicus Banks 1921 (AR1 Eastern west to TX NE MT)

(abdomen all dark)

            pepticus (Say) 1837 (CO ID IL IN KS MO NE NC TX WI)

(tarsi largely ferruginous, abdomen with four silvery fasciae, F 13-16mm)

            praedator W. Fox 1892 (MD to FL west to East TX)

(Uses Katydids; nocturnal excavators) (hind femur half to all red)

            sayi Banks 1942 (CO ID KS OK TX WY)

(abdominal 5 or 6 silvery, female hindtibia with slender spines)

            seminole Banks 1942 (FL only)

(hind femur nearly all red)

            validus Cresson 1873 (CO FL KS MI NE NC NY NJ TX WI)

(Uses Katydids; 1-5 per cell) (scape bright orange-red)

(head and thorax pale golden, abdomen with four silvery fasciae, legs yellow-ferruginous, F 18-22mm)

            werneri Bohart 1994 (AZ TX)


Subtribe Larrina


Larra (1 Nea) (Uses mole crickets; ground nesters; uses head and forelegs as scrapers; egg laid on cricket and released)

            analis Fabricius 1804 (AR2 FL GA KS NE NC TX)

(large deep black species with apex of abdomen bright red in female, male is all black and more pilose)

            bicolor Fabricius 1804 (FL)

            dives (Lepeletier) 1845 (NC)


Liris (16 Nea) (Uses Gryllidae; carries prey by antennae; constructs nest or borrows site; does not pack closure)

            argentatus (Beauvois) 1811 (AR1 CA CO CT FL IL KS MD NE NC NY PA TX VA WV)

(steely gray females with subhyaline wings)

            beatus (Cameron) 1889 (CA NC NM NY TX)

            muspa Pate 1943 (FL TX)

            panamensis muesebeki (Krombein) 1954 (FL only)

            partitus Krombein and Shanks 1984 (TX only)




Lyroda (2 Nea) (Uses Orthoptera; carried by antennae; serial feeders; ground nesters and preexisting burrows)

            subita Say 1837 (AR3 CO CT ID IN KS MD NY NC PA TX UT VA)

(Gryllids; favors genus Nemobius; 1-7 per cell)

(wings hyaline, apex dusky)

            triloba Say 1837 (AR1 CT KS TX)

(wings very dark fuscous)


Miscophus (15 Nea) (Uses spiders)

            americanus W. Fox 1890 (FL MD NE NJ NY NC PA VA) (3.5 to 4 mm males, apically infuscate wings)

            slossanae barberi (Krombein) 1954 (FL only)

            texanus (Ashmead) 1898 (TX only)


Plenoculus (15 Nea Mostly CA and SW) (Uses Miridae; nests in sandy soil; entrance open)

            cockerelli W. Fox 1894 (AZ CA NM TX)

            davisi W. Fox 1893 (AZ CA CO KS MO MT NE NY PA WI)

(mandibles except tip, tegulae, tibiae and tarsae more or less yellowish, abdomen with last segment reddish at tip)

            davisi atlanticus Viereck 1902 (CT NC)

            gillaspyi Krombein 1938 (TX only)


Solierella (49 Nea) (Uses Hemiptera; cavity nester and twig and stem nester; nests closed with pebbles and grass)

            foxii (Viereck) 1902 (NJ) (abdomen red)

            inermis (Cresson) 1873 (FL IL KS NC TX) (pronotum, postscutellum and all the tibiae creamy yellow, abdomen black)

            nigrans (Krombein) 1909 (CO WV)

            peckhami (Ashmead) 1897 (CA FL KS NY OR)

            plenoculoides (W. Fox) 1893 (CO GA SC NE NY WV) (Uses small grasshoppers)

            pulawskii R. Bohart 1991 (TX only)

            vierecki Rohwer 1909 (CA CO TX)




Belomicrus (40 Nea) (Uses bee flies and Myrids; nests in sandy soils)

            bridwelli Pate 1940 (FL TN VA)

            potawatomi Pate 1947 (IA)


Oxybelus (41 Nea) (Uses Dipterids esp. Muscoidea; prey impaled on sting or held by middle legs; nests in sandy soil; loose aggregations)

            bipunctatus Olivier 1812 (ME MD NY PA) (favors Stratiomyiidae; 2-12 per cell)

            crandalli Bohart and Schlinger 1956 (AZ AR)

            cressonii Robertson 1889 (MI NE NC PA TX UT VA)

            decorosus (Mickel) 1916 (NE PA WV)

            emarginatus Say 1837 (AR3 CA CO FL IL IN KS MD MA NC NM NY TX VA WA WY) (favors Milichiidae; 4-38 per cell)

            exclamans Viereck 1906 (CA FL KS NE NM)

            inornatus (Robertson) 1901 (IL NC PA)

            laetus Say 1837 (AR1 FL IL IN NC TX VA)

            laetus fulvipes Ch. Robertson 1889 (FL)

            major Mickel 1916 (NE NC)

            mexicanus Ch. Robertson 1889 (IL)

            niger Ch. Robertson 1889 (IL NY)

            packardii Ch. Robertson 1889 (AR2 IA IL NE NM NC TX)

            rancocas Pate 1943 (NJ)

            sericeus Ch. Robertson 1889 (CA FL IL MS NE NY NC) (favors Otitidae; 3-14 per cell)

            subcornutus Cockerell 1895 (CO NE NM NY PA TX)

            subulatus CH. Robertson 1867 (AR1 CO IL NE ND NY) (uses Therevidae; 3-11 per cell)

            uniglumis (Linnaeus) 1758 (CA CO ID IL KS MT NE NM NY PA UT) (favors Anthomyiidae; 2-13 per cell)




Trypoxylon (32 Nea) (Uses spiders; most are hollow twig nesters, also stems, keyholes, preexisting burrows; nectar on Aster and Goldenrod)

            albitarse Fabricius 1804 (NJ PA VA)

            bridwelli Sandhouse 1940 (TX only)

            carinatum Say 1837 (IN MD NC TX WV)

            clarkei Krombein 1962 (MD VA WI)

(favors Mangora in Krombein)

            clavatum Say 1837 (AR4 CO FL GA KS NE NC MD MO NY TN TX VA WV)

(Krombein found 80% "wandering spider" use including Crab Spiders and Jumpers; 5-22 per cell; males assist at nest)

            clavatum johannis Richards 1934 (FL GA)

            collinum F. Smith 1856 (FL GA)

            collinum rubrocinctum Packard 1867 (AR1 DE KS MO NC NE NY MD PA SC TN TX VA)

(males assist in nest protection; uses 100% orb weavers, mostly Araneidae and Theridion; 5-27 per cell; favored dead tree traps for Krombein; provisions in July and August in NY)

            fastigium W. Fox 1891 (NE NC TX)

            frigidum F. Smith 1856 (CA CO IA NE NM NY NC MO MD NV PA TX VA WV WI WY) (favors structural sites in Krombein)

            johnsoni W. Fox 1891 (DC FL NC NJ MD MO NY TN VA WV) (favors Tetragnathids in Krombein)

            lactitarse de Saussure 1867 (FL GA KS NE NC PA TN TX VA WV WI)

            pennsylvanicum de Saussure 1867 (ID MD NC NY PA VA WV WI)

            politum Say 1837 (AR2 FL IN LA MD NE NC SC TX VA WV)

(favors Araneidae; 2-17 per cell; pipe organ mud nests)

            richardsi Sandhouse 1940 (MD TX VA WV)

            spinosum Cameron 1889 (TX only)

            texense Saussure 1867 (AR2 FL CO IL KS NE NM TX)

            tridentatum Packard 1867 (AR1 AZ CA FL KS LA MD MO NE NC NJ NY NV UT VA WV)

(favors Araneidae, occasionally takes Black Widows, males assist at nest, Krombein found only orb weavers in nests; all one species in some studies; 7-12 per cell; prefers open areas)

            tridentatum archboldi Krombein 1959 (FL only)






Mellinus (4 Nea)

            bimaculatus Packard 1867 (CT IN NH NJ PA)




Pemphredonini (Uses Aphids)


Subtribe Pemphredonina


Passaloecus (16 Nea) (Uses aphids; twig and wood nester; plugged and partitioned with resin or pebbles; some species steal from other aphid feeding wasps)

            annulatus (Say) 1837 (IN KS NE NJ NC PA TX VA WV)

            areolatus Vincent 1979 (MD NY VA WV)

            cuspidatus F. Smith 1856 (CA CO DC MA NC NY VA WV) (14 -74 aphids per cell)

            lineatus Vincent 1979 (WV)

            relativus W. Fox 1892 (CO PA WY)

            singularis Dahlbom 1844 (NY)


Pemphredon (9 Nea) (Nests in tubes and bores and galls, decayed wood; nests are open; carries aphids in mandibles)

            foxii Rohwer 1917 (NC NJ VA WV)

            inornata Say 1824 (CO CT NE PA WI WV)

            lugubris (Fabricius) 1793 (AZ CO NC NH)

            menckei R. Bohart 1993 (MD)

            montana Dahlbom 1845 (CT CO NH NC)

            nearctica Kohl 1890 (CO NV WV)

            pulawskii Dollfuss 1993 (VA)

            unicolor Panzer 1798 (NJ TX WV)


Subtribe Spilomenina


Spilomena (5 Nea) (Uses immature and adult thrips; nests in twigs, timber, decayed wood)

            alboclypeata Bradley 1906 (WV WY)

            ampliceps Krombein 1952 (WV)

            barberi Krombein 1962 (CA MD PA VA)

            pusillus Say 1837 (CT IN MD NY NC VA WV)


Xysma (1 Nea) (Uses thrips; nests in Anobiid borings in sheds and buildings)

            ceanothae (Viereck) 1904 (DC MD PA)


Subtribe Stigmina


Stigmus (8 Nea) (Uses aphids; twig nesters; carries prey with mandibles; small active black wasps)

            americanus Packard 1867 (CO CT IL KS KY NC NY PA VA WV MY)

            aphidiperda Rohwer 1911 (NC PA)

            fraternus Say 1824 (CO MO PA VA WV)

            fulvicornis Rohwer 1923 (MS)

            fulvipes W. Fox 1892 (CA NE NM NC VA)

            inordinatus universitatus Rohwer 1909 (CO WV)


Psenini (Use hoppers)


Mimesa (33 Nea)

            cressoni Packard 1867 (CO IL IN IA MD MO NM NE NJ NH NY NC UT)

(9-17 items per cell) (second and third tergites red, petiole of abdomen slender, at least as long as hind femur)

            ezra Cresson 1865 (CO IL MD NY)

            lutaria (Fabricius) 1787 (GA ME MD NM NY NE)

            maculipes W. Fox 1893 (FL NC NJ)

(abdomen entirely without red basal markings, apices of tergites brownish-yellowish, petiole of abdomen thick and shorter than hind femur)

            pauper Packard 1867 (IL ME NE ND VA WV)

(apex of first and all of second tergite red)


Mimumesa (16 Nea) (Uses Homoptera, Delphacids and Cicadellids; nests in clay banks, decaying wood)

            canadensis (Malloch) 1933 (NY)

(antennal flagellum with entire lower side bright yellow)

            fuscipes (Packard) 1867 (MA)

            johnsoni (Viereck) 1901 (MD NJ PA VA WV)

(female with golden facial pubescence, males silver haired on face, rufous on basal third of antenna)

            leucopus (Say) 1837 (CO IL IN MD)

(face densely covered in female with oppressed silvery hairs)

            longicornis (W. Fox) 1898 (FL LA MD NC NJ SC)

            mellipes (Say) 1837 (IN IL IA NE NY OH)

(antennal flagellum with entire lower side bright yellow)

            nigra (Packard) 1867 (CT NC PA VA WV)

(antennal flagellum entirely dark)

            regularis (W. Fox) 1898 (NJ)


Pluto (21 Nea) (Uses leafhoppers; gregarious)

            aerofacies (Malloch) 1933 (NC TX)

            albifacies (Malloch) 1933 (IA TX)

            angulicornis (Malloch) 1933 (IA LA NM TX)

            arenivagus (Malloch) 1933 (FL GA NC)

            littoralis (Malloch) 1933 (FL MD NC SC)

            rotundus van Lith 1979 (FL)

            rufibasis (Malloch) 1933 (FL GA LA MD NC)

            sayi (Cresson) 1873 (AR2 CA KS MO NC TN TX VA)

            suffusus (W. Fox) 1898 (CA FL NM TX)

            tibialis (Cresson) 1873 (AL DC FL NC SC TX VA)


Psen (4 Nea) (Nests in stumps and dead trees, hard sand, clay banks)

            barthi Viereck 1907 (MD NC PA WI)

            erythropoda Rohwer 1910 (MD NY NC VA WV)

(petiole red or brownish, never black)

            monticola (Packard) 1867 (AL CT DC MD NH NC VA WV)

(bright red abdomen and black petiole)


Pseneo (3 Nea) (Uses leafhoppers; ground nesting)

            punctatus carolina (Rohwer) 1910 (NC)

            punctatus ferruginea (Viereck) 1901 (FL)

            simplicicornis (W. Fox) 1898 (NC)



Psenulus (4 Nea) (Uses aphids and leafhoppers; nests in hollow stems, abandoned beetle borings)

            trisulcus (W. Fox) 1898 (CT MO NH NC NY PA VA WV)






Aphilanthops (4 Nea) (uses various ant species)

            frigidus (F. Smith) 1856 2nd (AR1 CO CT IL MI MN NE NC ND NY) (2-3 per cell)


Clypeadon (8 Nea)

            dreisbachi (R. Bohart) 1959 (CO OK TX)


Cercerini Beetle Hunters


Cerceris (87 Nea) (Uses adult Coleoptera) (See characters for Scullen groups I-V here)

            alaope Banks 1912 (AR1 IL KS MA MO NJ NY NC VA)

(bands on tergum 2 deeply emarginate, black and yellow species, no amber or brown, close to clypeata)

            arelate Banks 1912 (IA MA ME MN NH NY NC PA VT VA)

(lateral apices of the clypeal process very acute and sides subparallel, related to nigrescens)

            astarte Banks 1913 (IL IN MA MI NC NH NY VA WI WV)

(pygidium sides subparallel, clypeal sides convergent, markings black and creamy yellow, no ferruginous or amber)

            atramontensis Banks 1913 (AR IL KS MD MA MI MN NY NC NE PA TX VA WI)

(band on tergum 2 broader than on other terga, black and yellow species)

            bicornuta Guerin Meneville 1844 2nd (AR2 Ext Central and Eastern)

(uses Billbugs) (clypeal process bicuspidate, without prolonged lateral processes, 16mm or longer females of black, ferruginous, yellow and brown, variable)

            blakei Cresson 1856 (FL GA IL KS LA NC TX)

(Group I females, scoop-shaped clypeus, female face largely reddish yellow, distinct red area covering one or more anterior abdominal terga)

(uses weevils and Chrysomelids)

            clypeata Dahlbom 1844 (AR7 Ext Central and Eastern)

(prey weevils, rarely Chrysomelids) (bands on terga 3 to 5 narrow but unbroken and broad laterally, band on tergum 2 broader than those on 3 to 5 with little or no emargination, black and yellow females, no amber or brown)

            compacta Cresson 1865 (AR6 Ext Central and Eastern)

(Group III females, band on tergum 2 broader than others, most terga deeply emarginate with black, propodeum with little or no yellow, metanotom yellow)

            compar Cresson 1865 (AZ FL GA IL IN IA KS MI NE NC OH OK PA TX VT VA)

(Group V females, subequal narrow bands on all terga)

            crucis Viereck & Cockerelli 1904 (AR1 CO NM TX NC NE VA)

(Group I females, bands on terga narrow and deeply emarginate, propodeum black and immaculate, occiput black except for small yellow spots at back of compound eyes, clypeal border with two widely separated hyaline points)

            dentifrons Cresson 1865 (IL IA MA MI MN ND NE NC NY PA WI)

(small 10mm females, lateral apices of the processes of the clypeus considerably prolonged, doubly indented band on tergum 1, deeply emarginate band on tergum 2)

            deserta Say 1824 (CO to MT west to NC and ME)

(bands on terga 2 to 5 subequal, 10 to 11mm females, clypeal process very short)

            echo Mickel 1916 (GA IL IA KS MA ME NE NJ NY NV NC TX VA WI)

(Group I females [see blakei and crucis], face is dark, has the distinct red area covering one or more abdominal terga)

            finitima Cresson 1865 (AR2 Ext Central and Eastern) (Group I females, not separated from crucis by Scullen)

            flavofasciata H. Smith 1908 (AL AR FL GA IL IA KS MO NE NC OK)

(uses Chrysomelids; 12-30 per cell) (sand lover) (Group III females, limited yellow markings, wide yellow band on distal half of tergum 2, no red or amber)

            frontata Say 1823 (AR1 AZ CO IL IA KS MO NM OK TX VT)

(very sexually dimorphic, in MO uses two weevil species, nests in very hard packed earth and roads) (clypeal process bicuspidate)

            fumipennis Say 1837 (AR2 FL IN KS MD MS MO MN NC NY TN TX)

(uses Buprestidae; rarely uses weevils; 5-18 per cell) (Group II females, one complete, wide ivory band on second tergum, wings black, ridged enclosure surface)

            halone Banks 1912 (AR2 IL IA MD MI MN MO NY NC ND NE OH PA SC TX VA WV WI)

(uses Curculionidae) (13 to 15mm females, band on tergum 2 much broader than on the other terga but deeply emarginate, clypeal process very short)

            insolita Cresson 1865 (AL AR4 AZ FL GA IL IA MD MN MS NC NE NM OH OK TX VA WV)

(Group V females [see compar], second tergum immaculate or with small lateral spots, hindfemur variable shades of amber)

            intricata graphica F. Smith 1873 (AZ IL KS NE NM OK TX)

(Group IV females, 15-18 mm, thorax largely reddish amber)

            irene Banks 1912 (LA MN NE OK TX)

(Group I females, no white markings, tegulae pitted and elevated, clypeal elevation a truncate cone)

            jucunda Cresson 1873 (FL IA IL KS KY NY NC OH TX)

(Scullen did not separate this species from compar)

            kennicottii Cresson 1865 (AL AR2 AZ FL GA IA IL KS LA MS MO NM NC OH TX VA)

(Group I females, apical border of clypeal process sinuate, abdomen black except for a small patch on tergum 1, deeply emarginate bands on terga 2 and 4 and two lateral patches on 3)

            mandibularis Patton 1880 (AL CT GA IL IA LA MD NE NJ NC PA TN VA WV)

(Group III females, tergum 1 with a small divided patch of yellow)

            nigrescens F. Smith 1856 (Ext Western including KS TX, MN and IL east to NC and ME)

(favors Curculionidae) (clypeal process convex above, sides subparallel)

            nitoides G. Ferguson 1984 (IL MI NY NC OH TN TX) (described after Scullen's paper)

            prominens Banks 1912 (GA IL IA KS LA MA MN NC PA VA)

(close to clypeata, band on tergum 2 deeply emarginate)

            rufopicta F. Smith 1856 (AR3 CT FL IL IA MI MN NE NC OH OK TX)

(Group III females, Scullen had divided into multiple subspecies under robertsonii, clypeal process lamellate)

            rufinoda Cresson 1865 (AR1 CO KS LA NC NE OH TX VA)

(Group I females, distinct red over one or more anterior abdominal terga)

            sexta Say 1837 (CO IL IN MA MI MO KS NE NY NC ND TX VA WI)

(15mm females, tergum 1 largely yellow)

            squamulifera Mickel 1916 (AR KS NE NC OK TX)

(Group III females, thorax largely black or amber yellow, tergal bands on all abdominal segments all yellow)

            tolteca de Saussure 1867 (AR1 AZ FL LA OK TX)

(tergum 1 with a broad medial yellow patch, tergum 2 black or dark amber without a yellow band)

            verticalis F. Smith 1856 (FL GA KS LA NC NE OK SC TX)

(large females 16mm or longer, pygidium narrowing basally to an acute or very narrow end)


Eucerceris (26 Nea 2)

            canaliculata (Say) 1823 (AR CO KS NM TX UT)

            zonata (Say) 1823 (AR KS MA NE NM TX)


Philanthini (Bee Wolves)


Subtribe Philanthina


Philanthus (32 Nea)

            albopilosus Cresson 1865 (CO IL IA MI NE ND OH)

            bilunatus Cresson 1865 (CO IL MI NM NY NC QU)

(Uses mostly Halictid bees, prefers firm coarse soil in wooded areas, less colonial)

            gibbosus (Fabricius) 1775 Live (AR4 CT FL IL IN KS MA MD MI MO NE NY NC PA TX UT)

(Uses mostly Halictid bees, colonies small and loose)

            lepidus Cresson 1865 (CO MD MI MN NY NC PA TX)

(Uses mostly Halictid bees, tight colonies in sparsely vegetated fine-grained sand)

            politus Say 1824 Live (FL IL MI MN MO NY NC PA TX)

(Uses mostly Halictids, rarely colonial, gathers then sorts bees)

            sanbornii Cresson 1865 (FL KS MA MN NE NC)

(Uses six or seven bee and wasp families, large colonies, one nest per season, flies mainly in June)

            solivagus Say 1837 (CT IN MI NE NY ND)

            ventilabris Fabricius 1798 (AR4 CA CO FL IL KS MI MN NY NC TX)

(Uses Halictid bees and wasps; 3-15 per cell; more packed soils, shallower nests, closes nests when away)


80 Genera

462 Species

12 Additional subspecies